Cao Kunfang 曹坤芳
14th Apr


Cao Kunfang 曹坤芳

KunfangResearch profile:

After joining Guangxi University in 2013, I have set up a new research group within the College of Forestry in Nanning, which focuses on the interplay between plant ecophysiology and plant evolution. My main research interests include the ecophysiology of tropical and subtropical plants in rainforests, dry forests and savanna systems, ecophysiology of economically important plants. In our projects we study water relations and hydraulics of plants, photosynthetic and photoprotection responses to stressful conditions/environments such as drought, cold spells and salinity in mangroves, evolution of plant functional traits and plant ecological adaptations.


Current Research Projects:

2017-2021, A major research grant for Distinguished Scholar of Guangxi Zhuang-Nationality Autonomous Region. (Principle Investigator)

2017-2020,Hydraulic structure and function and their coordination with stomatal regulation behavior. Funded by National Natural Science Foundation of China. (Principal Investigator)

2015-2018, Hydraulic structure and function of stems and leaves of basal angiosperms and their correlation with photosynthetic capacity. Funded by National Natural Science Foundation of China. (Principal Investigator)

2016-2019,Monitoring of dynamics of representative primary forests in Guangxi Zhuang-Nationality Autounomous Region. Funded by Guangxi Department of Science and Technology. (Principal Investigator)

2016-2020, Spatial pattern of plant diversity in Southwest China, a subproject under a major project funded by Ministry of Sciences and Technology of China. (Principal Investigator)

2013-2017 – Ecophysiological adaptation of tropical and subtropical plants to environmental stress, focusing on mangrove plants and basal angiosperms, by Guangxi University. (Principal Investigator)


Curriculum Vitae:


  • 1/1995 PhD, Forest Ecology and Silviculture, Wageningen Agricultural University, The Netherlands (supervisor: Prof. Dr. Ir. R.A.A. Oldeman)
  • 7/1982 B.Sc., Fruit Horticulture, Nanjing Agricultural University, China.

Other training

  • 1988-1989 Exchange study at Wageningen Agricultural University, the Netherlands (18 months)


  • Since 1/2013, Professor, Honorable Dean of College of Forestry, Guangxi University
  • 2006.04-2006.09, Visiting professor to Weed Science Center, Utsunomiya University, Japan.
  • 10/1999-12/2012, Professor and head of the Research Group for Physiological Plant Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences
  • 2-10/1999 Associate Professor at Institute of Botany, CAS.
  • 1997-1998 Research fellow at Universiti Brunei Darussalam, Brunei, 18 months.
  • 1995-1997 Postdoctoral fellow of Japan Society for Promotion of Science at Utsunomiya University, 2 years.
  • 1991-1994 Postgraduate researcher at Wageningen Agricultural University, Netherlands.
  • 1982-1990 Research Fellow at Institute of Botany, CAS.


  • 2016 Distinguished Scholar of Guangxi Zhuang-Nationality Autonomous Region
  • 2012 Excellent scientist, awarded by the State Council of China
  • 2009 Excellent teacher, awarded by Chinese Academy of Sciences
  • 2006 Outstanding Scientist Award by the Provincial Government of Yunnan.
  • 2003 National Award for Achievements of Overseas Returned Scholars, jointly awarded by the Chinese Ministry of Education, Ministry of Science and Technology, and Ministry of Personnel.

Members of Editorial Boards

  • Since 2017 Associate Editor, Journal of Ecology
  • 2010 – 2016      Associate Editor, Plant Ecology
  • 2003 – 2007      Forest Ecology and Mangement
  • 1999 – present  Journal of Plant Ecology (Formerly Acta Phytoecologia Sinica)
  • 1999 – present  Biodiversity Science (a national journal)
  • Since 2010       Wuhan Botanical Research (a national journal)

Reviewer for journals
Science, Ecology Letters, Plant Physiology, Ecology, New Phytologist, Plant, Cell and Environment, Tree Physiology, American Journal of Botany, Annals of Botany, Journal of Tropical Ecology, Acta Oecologia, Forest Ecology and Mangement, Annals of Forest Science, China Science,Biotropica, Ecological Research, Journal of Intergrated Plant Biology (formally Acta Botanica Sinica), Journal of Plant Ecology, Biodiversity Science, Acta Ecologia Sinica, Chinese Journal of Applied Ecology, Chinese Journal of Forestry Science, Jounal of Plant Science, Journal of Plant Taxomony and Resources, Journal of Mountain Science.

Professional Membership

  • British Society of Ecology
  • International Society of Ecology
  • International Association for Vegetation Science
  • Botanical Society of China, Forestry Society of China, Ecological Society of China

Representative publications in peer-reviewed (SCI) journals (*, author for correspondence)

– Wan-Li Zhao, Zafar Siddiq, Pei-Li Fu, Jiao-Lin Zhang, Kun-Fang Cao*. 2017. The stable stomatal number per minor vein length can indicate leaf water balance. Scientific Reports 7: 2211.
– Yong-Jiang Zhang#* , Lawren Sack# , Kun-Fang Cao# , Xue-Mei Wei & Nan Li, 2017. Speed versus endurance tradeoff in plants: Leaves with higher photosynthetic rates show stronger seasonal declines. Scientific Reports 7:42085, DOI: 10.1038/srep42085 (#, equal contribution author)
– Zhang, Shu-Bin, Jiao-Lin Zhang, and Kun-Fang Cao 2017. Divergent hydraulic safety strategies in three co-occurring Anacardiaceae tree species in a Chinese savanna. Frontiers in Plant Science 7, article 2075.
– Jiang, Guo-feng#, Uromi M. Goodale#, Yan-yan Liu, Guang-you Hao, Kun-fang Cao*, 2017. Salt management strategy defines the stem and leaf hydraulic characteristics of six mangrove tree species. Tree Physiology 37:389–401
– Ya-Jun Chen, Stefan A. Schnitzer, Yong-Jiang Zhang, Ze-Xin Fan, Guillermo Goldstein , Kyle W. Tomlinson, Hua Lin, Jiao-Lin Zhang* and Kun-Fang Cao*. 2017. Physiological regulation and efficient xylem water transport regulate diurnal water and carbon balances of tropical lianas. Functional Ecology 31:306–317
– Huang, Wei ; Yang, Ying-Jie; Hu, Hong, Cao, Kun-Fang; Zhang, Shi-Bao, 2016. Sustained Diurnal Stimulation of Cyclic Electron Flow in Two Tropical Tree Species Erythrophleum guineense and Khaya ivorensis, FRONTIERS IN PLANT SCIENCE 6: article 1068 (DOI: 10.3389/fpls.2015.01068)
– Gbadamassi G. O. Dossa, Ekananda Paudel, Kunfang Cao, Douglas Schaefer, Rhett D. Harrison, 2016. Factors controlling bark decomposition and its role in wood decomposition in five tropical tree species. Scientific Reports 6:34153.
– Wan-Li Zhao, Ya-Jun Chen, Timothy J. Brodribb, Kun-Fang Cao*. Weak co-ordination between vein and stomatal densities in 105 angiosperm tree species along altitudinal gradients in Southwest China. Functional Plant Biology 43:1126-1133.
– Yong-Jiang Zhang, Sandra J. Bucci, Nadia S. Arias, Fabian G. Scholz, Guang-You Hao, Kun-Fang Cao and Guillermo Goldstein, 2016. Freezing resistance in Patagonian woody shrubs: the role of cell wall elasticity and stem vessel size. Tree Physiology 36:1007-1018.
– Wei Huang*, Ying-Jie Yang, Hong Hu, Shi-Bao Zhang, Kun-Fang Cao, 2016. Evidence for the role of cyclic electron flow in photoprotection for oxygen-evolving complex. Journal of Plant Physiology 194:54-60.
– Shi-Dan Zhu#, Hui Liu#, Qiu-Yuan Xu, Kun-Fang Cao, Qing Ye*. 2016. Are leaves more vulnerable to cavitation than branches? Functional Ecology 30:1740-1744.
– Catherine Roumet, Marine Birouste, Catherine Picon-Cochard;,Murielle Ghestem, Normaniza Osman, Sylvain Vrignon-Brenas, Kun-fang Cao, Alexia Stokes. 2016. Root structure – function relationships in 74 species: evidence of a root economics spectrum related to carbon economy. New Phytologist 210: 815–826
– Pei-Li Fu, Wen-Jie Liu, Ze-Xin Fan and Kun-Fang Cao* 2016. Is fog an important water source for woody plants in an Asian tropical karst forest during dry season? Ecohydrology 9:964-972 (doi: 10.1002/eco.1694).
– Ya-Jun Chen*, Frans Bongers, Kyle Tomlinson, Ze-Xin Fan, Hua Lin, Shu-Bin Zhang, Yu-Long Zheng, Yang-Ping Li, Kun-Fang Cao*, and Jiao-Lin Zhang. 2016. Time lags between crown and basal sap flows in tropical lianas and co-occurring trees. Tree Physiology 36:736-747
– Bartlett, M. K., Y. Zhang, J. Yang, N. Kreidler, S-w. Sun, L. Lin, Y-H. Hu, K-F. Cao, L. Sack. 2016. Drought tolerance as a driver of tropical forest assembly: resolving spatial signatures for multiple processes. Ecology 97 (2) 503-514
– Zafar Siddiq and Kun-Fang Cao*, 2016. Increased water use in dry season in eight dipterocarp species in a common plantation in the northern boundary of Asian tropics. Ecohydrology 9:871-881.
– Sean M. Gleason, Mark Westoby, Steven Jansen, Brendan Choat, Uwe G. Hacke, Robert B. Pratt, Radika Bhaskar, Tim J. Brodribb, Sandra J. Bucci, Kun-Fang Cao, et al. 2016. Weak tradeoff between xylem safety and xylem-specific hydraulic efficiency across the world’s woody plant species. New Phytologist 209 (1): 125–138
– Ren-Yi Ma, Jiao-Lin Zhang, Molly A. Cavaleri, Frank Sterck, Joeri S. Strijk, Kun-Fang Cao*, 2015. Convergent evolution towards high net carbon gain efficiency contributes to the shade tolerance of palms (Arecaceae). Plos One 10(10): e0140384.
– Jun-Wen Chen and Kun-Fang Cao, 2015. A possible link between hydraulic properties and leaf habits in Hevea brasiliensis. Functional Plant Biology 42(8): 718-726.
– Gbadamassi G. O. Dossa, Ekananda Paudel, Hongyan Wang, Kunfang Cao, Douglas Schaefer, Rhett D. Harrison, 2015. Correct calculation of CO2 efflux using a closed-chamber linked to a non-dispersive infrared gas analyzer. Methods in Ecology and Evolution: 6(12):1435-1442
– Shi-Jian Yang*, Yong-Jiang Zhang, Guillermo Goldstein, Mei Sun, Ren-Yi Ma and Kun-Fang Cao*, 2015. Determinants of water circulation in a woody bamboo species: afternoon use and night-time recharge of culm water storage. Tree Physiology: 35(9):964-974
– Guofang Liu, William K. Cornwell, Kunfang Cao,Yukun Hu, Richardus S.P. Van Logtestijn; Shijian Yang, Xiufang Xie, Yalin Zhang, Duo Ye, Xu Pan, Xuehua Ye, Zhenying Huang, Ming Dong, Johannes H.C. Cornelissen, 2015. Termites amplify effects of wood traits on decomposition rates among multiple bamboo and dicot woody species. Journal of Ecology 103 (5): 1214–1223
– Yong-Jiang Zhang*, Kun-Fang Cao*, Lawren Sack, Nan Li, Xue-Mei Wei and Guillermo Goldstein, 2015. Extending the generality of leaf economic design principles in the cycads, an ancient lineage. New Phytologist 206: 817–829.
– Shi-Dan Zhu, Ya-Jun Chen, Kun-Fang Cao, Qing Ye, 2015. Interspecific variation in branch and leaf traits among three Syzygium tree species from different successional tropical forests. Functional Plant Biology 42: 423–432.
– Chen, Ya-Jun#, Kun-Fang Cao# , Stefan A. Schnitzer, Ze-Xin Fan, Jiao-Lin Zhang, Frans Bongers, 2015. Water-use advantage for lianas over trees in tropical seasonal forests. New Phytologist 205 (1): 128–136.
– Yong-Jiang Zhang*, N. Michele Holbrook, and Kun-Fang Cao* 2014. Seasonal dynamics in photosynthesis of woody plants at the northern limit of Asian tropics: potential role of fog in maintaining tropical rainforests and agricultures in Southwest China. Tree Physiology 34(10): 1069–1078.
– Tiange Lang, Kangquan Yin, Jinyu Liu, Kunfang Cao, Charles H. Cannon, Fang K. Du. 2014. Protein Domain Analysis of Genomic Sequence Data Reveals Regulation of LRR Related Domains in Plant Transpiration in Ficus. PLOS ONE 9(9): e108719 IF=3.534
– Megan K. Bartlett, Ya Zhang, Nissa Kreidler, Shanwen Sun, Rico Ardy, Kunfang Cao, and Lawren Sack 2014. Global analysis of plasticity in turgor loss point, a key drought tolerance trait. Ecology Letters 17(12): 1580–1590. IF(2013)=13.042. (I区)
– Murielle Ghestem, Kunfang Cao, Wenzhang Ma, Nick Rowe, Raphaëlle Leclerc, Clément Gadenne, Alexia Stokes 2014. A framework for identifying plant species to be used as ‘ecological engineers’ for fixing soil on unstable slopes. Plos One 9(8): e95876.
– Chen, Ya-Jun, Frans Bongers, Jiao-Lin Zhang, Jin-Yu Liu, & Kun-Fang Cao*, 2014. Different biomechanical design and ecophysiological strategies in juveniles of two lianas species of Rhamnaceae with contrasting growth habit. American Journal of Botany 101(6): 925–934.
– Guofang Liu, William K. Cornwell, Xu Pan, Kunfang Cao, Xuehua Ye, Zhenying Huang, Ming Dong and Johannes H. C. Cornelissen 2014. Understanding the ecosystem implications of the angiosperm rise to dominance: leaf litter decomposability among magnoliids and other basal angiosperms. Journal of Ecology 2014, 102, 337–344
– Li, Shuai, Yong-Jiang Zhang, Lawren Sack, Christine Scoffoni, Atsushi Ishida, Ya-Jun Chen, Kun-Fang Cao*, 2013. The heterogeneity and spatial patterning of structure and physiology across the leaf surface in giant leaves of Alocasia macrorrhiza. PLoS One 8(6):e66016.
– Zhang,Yong-Jiang, Qiu-Yun Yang, David W. Lee, Guillermo Goldstein, Kun-Fang Cao*, 2013. Extended leaf senescence promotes carbon gain and nutrient resorption: importance of maintaining winter photosynthesis in subtropical forests. Oecologia (in press, on-line).
– Hao, Guangyou; Wang, Ai-Ying; Sack, Lawren; Goldstein, Guillermo ; Cao, Kun-Fang*, 2013. Is hemiepiphytism an adaptation to high irradiance? Testing seedling responses to light levels and drought in hemiepiphytic and non-hemiepiphytic Ficus. Physiologia Plantarum 148: 74–86.
– Huang, Wei, Pei-Li Fu, Yan-Juan Jiang, Jiao-Lin Zhang, Shi-Bao Zhang, Hong Hu* and Kun-Fang Cao*, 2013.Differences in the responses of photosystem I and photosystem II of three tree species Cleistanthus sumatranus, Celtis philippensis and Pistacia weinmannifolia exposed to a prolonged drought in a tropical limestone forest.Tree Physiology 33:211-220.
– Zhang,Shi-Bao, Kun-Fang Cao*, Ze-Xin Fan, Jiao-Lin Zhang, 2013. Potential hydraulic efficiency in angiosperm trees increases with growth-site temperature but has no trade-off with mechanical strength. Global Ecology and Biogeography 22, 971–981
– Zhang, Yong-Jiang, F.C. Meinzer, Jin-Hua Qi, G. Goldstein, Kun-Fang Cao*, 2013. Midday stomatal conductance is more related to stem rather than leaf water status in subtropical deciduous and evergreen broadleaf trees. Plant, Cell and Environment 36, 149–158.
– Zhang,Yong-Jiang, Qiu-Yun Yang, David W. Lee, Guillermo Goldstein, Kun-Fang Cao*, 2013. Extended leaf senescence promotes carbon gain and nutrient resorption: importance of maintaining winter photosynthesis in subtropical forests. Oecologia 173:721-730.
– Tan, Zheng-Hong, Yi-Ping Zhang, Naishen Liang, Yue-Joe Hsia, Yong-JiangZhang, Guo-Yi Zhou, Yue-Lin Li, Jehn-Yih Juang, Hou-Sen Chu, Jun-HuaYan, Gui-Rui Yu, Xiao-Min Sun, Qin-Hai Song, Kun-Fang Cao, D.A. Schaefer and Yu-Hong Liu, 2012. An observational study of the carbon-sink strength of East Asian subtropical evergreen forests. Environmental Research Letters 7:044017(12pp).
– Bartlett, Megan, Christine Scoffoni, Rico Ardy, Ya Zhang, Shanwen Sun, Kunfang Cao, Lawren Sack, 2012. Rapid determination of comparative drought tolerance traits: using an osmometer to predict turgor loss point. Methods in Ecology and Evolution 3(5): 880-888
– Zhang, Jiao-lin, L. Poorter · Kun-Fang Cao*, 2012. Photosynthetic thermotolerance is correlated with leaf lifespan across woody savanna species. Annuals of Botany 110: 1027–1033 ( IF=4.03)
– Cao,Kun-Fang,Shi-Jian Yang,Yong-jiang Zhang,T.J. Brodribb, 2012. Maximum height of grasses is determined by roots. Ecology Letters 15:666-672.
– Fu, Pei-Li, Yan-Juan Jiang, Ai-Ying Wang, Tim J. Brodribb, Jiao-Lin Zhang, Shi-Dan Zhu, Kun-Fang Cao*, 2012. Stem hydraulic traits and leaf water-stress tolerance are coordinated with the leaf phenology of angiosperm trees in an Asian tropical dry karst forest. Annual of Botany 110:189-199.
– Fan, Ze-Xin, Shi-Bao Zhang, Guang-You Hao, J.W. Ferry Slik, Kun-Fang Cao*. 2012. Hydraulic conductivity traits predict growth rates and adult stature of 40 Asian tropical tree species better than wood density. Journal of Ecology 100:732-741.
– Huang, Wei, Shi-Bao Zhang, Kun-Fang Cao* 2012. Evidence for leaf fold to remedy the deficiency of physiological photoprotection for photosystem II. Photosynthesis Research 110: 185-191.
– Huang, Wei, Shi-Jian Yang, Shi-Bao Zhang, Jiao-Lin Zhang, Kun-Fang Cao*. 2012. Cyclic electron flow plays an important role in photoprotection for the resurrection plant Paraboea rufescens under drought stress. Planta 235 (4):819-828.
– Zhang, Shi-Bao, Jiao-Lin Zhang, Ferry Slik & Kun-Fang Cao*. 2012. Leaf element concentrations of terrestrial plants across China are influenced by taxonomy and the environment. Global Ecology and Biogeography 21:809–818.
– Yang, Shi-jian, Yong-jiang Zhang, Kun-Fang Cao*, 2012. Recovery of diurnal depression of leaf hydraulic conductance in a subtropical woody bamboo species: embolism refilling by nocturnal root pressure. Tree Physiology 32: 414–422.
– Chen, Jun-Wen, Qiang Zhang, Xiao-Shuang Li and Kun-Fang Cao, 2011. Steady and dynamic photosynthetic responses of seedlings from contrasting successional groups under low light growth conditions. Physiologia Plantarum 141:84-95.
– Fan, Ze-Xin, Achim Bräuning, Axel Thomas, Jin-Bao Lid and Kun-Fang Cao. 2011. Spatial and temporal temperature trends on the Yunnan Plateau (Southwest China) during 1961–2004. International Journal of Climatology 31: 2078-2090.
– Hao, Guang-You, Guillermo Goldstein, Lawren Sack, N. Michele Holbrook, Zhi-Hui Liu, Ai-Ying Wang, Rhett D. Harrison, Zhi-Hui Su, Kun-Fang Cao. 2011. Ecology of hemi-epiphytism in fig species is based on evolutionary correlation of hydraulics and carbon economy. Ecology 92 (11): 2117-2130.
– Hao,Guang-You, Ai-Ying Wang, Zhi-Hui Liu, Augusto C. Franco, Guillermo Goldstein and Kun-Fang Cao*, 2011. Differentiation in light energy dissipation between hemiepiphytic and non-hemiepiphytic Ficus species with contrasting xylem hydraulic conductivity. Tree Physiology 31(6): 626-636 (the cover paper).
– Huang, Wei, Shi-Bao Zhang and Kun-Fang Cao*. 2011. Cyclic electron flow plays an important role in photoprotection for tropical trees illuminated at temporal chilling temperature. Plant and Cell Physiology 52(2): 297–305.
– Zhang,Shi-Bao, Zhi-Jie Guan, Wei Chang, Hong Hu, Kun-Fang Cao. 2011. Slow photosynthetic induction and low photosynthesis in Paphiopedilum armeniacum is related to its lack of guard cell chloroplast and peculiar stomatal anatomy. Physiologia Plantarum 142: 118-127.
– Zhang SB, F Slik, JL Zhang & KF Cao*. 2011. Spatial patterns of wood traits in China are controlled by phylogeny and environment. Global Ecology and Biogeography 20: 241–250.
– Huang, Wei, Shi-Bao Zhang and Kun-Fang Cao* 2010. Stimulation of cyclic electron flow during recovery after chilling-induced photoinhibition of photosystem II. Plant and Cell Physiology: 51(11): 1922-1928.
– Fan, Ze-Xin, Achim Bräuning, Qin-Hua Tian, Bao Yang, Kun-Fang Cao. 2010. Tree ring recorded May–August temperature variations since A.D. 1585 in the Gaoligong Mountains, southeastern Tibetan Plateau. Palaeogeography, Palaeoclimatology, Palaeoecology 296: 94–102.
– Zhu, Shi-Dan and Kun-Fang Cao*. 2010. Contrasting cost-benefit strategy between lianas and trees in a tropical seasonal rain forest in Southwest China. Oecologia 163:591–599.
– Hao, Guang-You, Lawren Sack, Ai-Ying Wang, Kun-Fang Cao* and Guillermo Goldstein*. 2010. Differentiation of water flux and drought tolerance traits in leaves of hemiepiphytic and non-hemiepiphytic Ficus species. Functional Ecology 24:731–740
– Wei Huang, Shi-Bao Zhang and Kun-Fang Cao*. 2010. The different effects of chilling stress under moderate illumination on photosystem II compared with photosystem I and subsequent recovery in tropical tree species. Photosynthesis Research 103:175–182.
– Chen, Jun-Wen, Qiang Zhang and Kun-Fang Cao, 2010. Gas exchange and hydraulics in seedlings of Hevea brasiliensis during water stress and recovery. Tree Physiology 30, 876–885.
– Zheng, Li; Ives, Anthony*; Garland, Theodore; Larget, Bret; Yu, Yang; Cao, Kun-fang*. 2009. New multivariate tests for phylogenetic signal and trait correlations applied to ecophysiological phenotypes of nine Manglietia species. Functional Ecology 23: 1059–1069.
– Zhang, Yong-jiang, FC Meinzer, Guang-you Hao FG Scholz, SJ Bucci, AC Franco, R Villalobos-Vega, JP Giraldo, Kun-fang Cao, WA Hoffmann, & G. Goldstein. 2009. Size-dependent mortality in a Neotropical savanna tree: the role of height-related adjustments in hydraulic architecture and carbon allocation. Plant, Cell and Environment 32:1456–1466.
– Chen, Jun-Wen, Qiang Zhang, Xiao-Shuang Li and Kun-Fang Cao*. 2009. Independence of stem and leaf hydraulic traits in six Euphorbiaceae tree species with contrasting leaf phenology. Planta 230:459–468.
– Fan, Ze-Xin, Achim Bräuning, Kun-Fang Cao & Shi-Dan Zhu. 2009. Growth-climate responses of high-elevation conifers in the central Hengduan Mountains, southwestern China. Forest Ecology and Management 258: 306–313
– Hao, G-Y, T.J. Jones, C. Luton, Y.-J. Zhang, E. Manzane, F.G. Scholz, S. J. Bucci, K.-F. Cao and G. Goldstein. 2009. Hydraulic redistribution in dwarf Rhizophora mangle trees driven by interstitial soil water salinity gradients: impacts on hydraulic architecture and gas exchange. Tree Physiology 29(5): 697-705.
– Zhang, Jiao-Lin and Kun-Fang Cao*. 2009. Stem hydraulics mediates leaf water status, carbon gain, nutrient use efficiencies and plant growth rates across dipterocarp species. Functional Ecology 23: 658–667
– Fan, Ze-Xin, Achim Bräuning, Bao Yang, Kun-Fang Cao. 2009. Tree ring density-based warm-season temperature reconstruction for the central Hengduan Mountains in southern China. Global and Planetary Change 65: 1–11.
– Zhang J.-L. L.-Z. Meng and K.-F. Cao*. 2009. Sustained diurnal photosynthetic depression in upmost canopy leaves of four dipterocarp species in the rainy and dry seasons: does photorespiration play a role in photoprotection? Tree Physiology 29: 217–228.
– Zhu, Jun-Jie, Jiao-Lin Zhang, Hong-Cheng Liu and Kun-Fang Cao*. 2009. Photosynthesis, non-photochemical pathways and activities of antioxidant enzymes in a resilient evergreen oak under different climatic conditions from a valley-savanna in Southwest China. Physiologia Plantarum 135: 62–72.
– Fan, Ze-Xin, Achim Bräuning & Kun-Fang Cao. 2008. Tree-ring based drought reconstruction in the central Hengduan Mountain region (China) since A.D. 1655. International Journal of Climatology 28:1879-1887.
– Hao, G-Y.,W. A. Hoffmann, F. G. Scholz, S. J. Bucci, F. C. Meinzer, A. C. Franco, K-F Cao and G. Goldstein. 2008. Stem and leaf hydraulics of congeneric tree species from adjacent tropical savanna and forest ecosystems. Oecologia 155: 405–415.
– Cai, Z-Q., L. Poorter, K.-F. Cao & F. Bongers. 2007. Seedling growth strategies in Bauhinia species: comparing lianas and trees. Annuals of Botany 100:831-838.
Wilske, B., K.-F. Cao, G. Schebeske, J.-W. Chen, A. Wang, J. Kesselmeier 2007. Isoprenoid emissions of trees in a tropical rainforest in Xishuangbanna, SW China. Atmospheric Environment 41: 3748–3757.
– Cao, Kun-fang, 2001. Morphology and growth of evergreen and deciduous broad-leaved saplings in a Chinese beech forests with dense bamboo undergrowth. Ecological Research 16 (3):509-517.
– Cao, Kun-fang & Booth, W.E.2001. Leaf anatomical structure and photosynthetic induction for seedlings of five dipterocarp species under contrasting light conditions in a Bornean heath forest. Journal of Tropical Ecology 17: 163-176
– Cao, Kun-fang 2000. Leaf anatomy and chlorophyll content of twelve woody species growing in contrasting light conditions in a Bornean heath forest. Canadian Journal of Botany 78:1245-1253
– Cao, Kun-fang 2000. Water relations and gas exchange of tropical saplings during a prolonged drought in a Bornean heath forest, with reference to root architecture. Journal of Tropical Ecology 16:101-116.)
– Cao, Kun-fang & Ohkubo, T. 1999. Suppression and release during canopy recruitment in Fagus crenata and Acer mono in two old-growth forests in Japan. Plant Ecology 145: 281-290.
– Cao, Kun-fang & R. Peters. 1998. Structure and stem growth of multi-stemmed trees of Fagus engleriana in China. Plant Ecology 139: 211-220.
– Cao, Kun-fang, Ohkubo, T., 1998. Allometry, root/shoot ratio and root architecture in understorey saplings of deciduous dicotyledonous trees in central Japan. Ecological Research 13: 217-227.
– Cao, Kun-fang, Peters, R. 1997. Species diversity of Chinese beech forests in relation to warmth and climatic disturbances. Ecological Research 12:175-189.
– Cao, Kun-fang, Peters, R. & Oldeman, R.A.A. 1995. Climatic range and distribution of Chinese Fagus species. Journal of Vegetation Science 6: 317-324.


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